Grasses and grass-like plants
Grasses and grass-like plants; graminoides.
Poaceae/Gramineae -> grass family
Cyperaceae -> sedge family
Juncaceae -> rush family
Poaceae/Gramineae -> grass family
Contains about 8,500 species in approximatly 650
genera
Ecomomically important
Cereal grains: wheat; corn; rice; barley; rye; etc.
Forage for livestock
Turf for lawns; parks; athletic fields
Source of cover and food for wildlife
Source of construction materials and utensils
Vegetative grass structure
Roots: adventitious and fibrous
Stems: specialized types: rhizomes;
stolons; runners
Culm is the upright aerial stem
Recognition Characteristics:
Jointed with swollen nodes; often
hollow at the internodes
Circular or ellipsoid in cross-section
Leaves: 2-ranked
Leaf consists of the blade; open
sheath with ligule; auricles may be present
on the blade; veination is parallel.
Reproductive structure
Inflorescence -> a spikelet which is a two-rankes cluster of florets (small flowers) typically subtended by a pair of glumes (sterile bracts).
Grass flower: adapted for wind or self pollination
Lemma
Palea
Locicules
Rachilla
Awns
Sterile lemma
Sterile floret
Stamens: 3 (1-6 or numerous)
Carpels 3 (often appearing as
2) stigmas 2 (-3) plumose
Grass fruit: a grain or caryopsis
Examples of grasses that typify variation within the family members found in the Gulf Coast Region:
Arundinaria; Hydrochloa; Monanthochloe; Erianthus;
Schizachrium; Andropogon; Phragmites; Ctenium; Spartina; Paspalum; Panicum;
Zizaniopsis; Setaria; Manisuris; Chinochloa; Eragrostis; Briza; Aristida;
Muhlenbergia.
Grass identification literature:
Chase, A. 1950. Manual of the Grasses of the United States by A. S. Hetchcock. U.S.D.A. Miscellaneous Publication No. 200. U.S. Government Printing Office, Washington.
Godfrey, R. K. and J. W. Wooten. 1979. Aquatic and Wetland Plants of Southeastern United States; Monocotyledons. University of Georgia Press, Athens. [ISBN: 0-8203-0420-4; LC card number QK125.G6, 584.0976, 76-28924].
Gould, F. W. 1975. The Grasses of Texas. Texas A & M University Press, College Station. [ISBN 0-89096-005-4, LC card number: QK495.G74G726, 584.9097641 75-18688].
Hatch, S. L., J. L. Schuster, and D. L. Drawe. 1999. Grasses of the Texas Gulf Prairies and Marshes. Texas A & M University Press, College Station. [ISBN 0-89096-875-6; LC card number: QK495.G74H34365, 1999, 584.909764-dc21].
Stutzenbaker, C. D. 1999. Aquatic and Wetland Plants
of the Western Gulf Coast. Texas Parks and Wildlife Press. [ISBN 1-885696-31-0].
Juncaceae -> Rush family
Worldwide: 6 genera, 400 species. Louisiana: 2 genera,
30 species.
Juncus and Luzula are two genera found
in our region
Juncus: glabrous plants; capsules with many
powderlike seeds
Luzula: hairy plants; capsules each with
3 seeds.
Vegetative features:
Herbs
Stems: round and solid, not jointed.
Leaves: 3-ranked, basal or along lower portion of
the stem
Composed of a blade and a usually open sheath lacking
a ligule
Blade linear, grass-like, may be reduced or absent
Reproductive features
Inflorescence highly branched, but often condensed
and head-like
Inflorescence bract may be upright and appear as
a continuation of the stem giving the appearance that the flowers are growing
from one side of the stem.
i.e.: J. effusus; J. romerianus;
Flowers: 6 distinct perianth (tepals) parts, scale-like.
Stamens: (3-) 6
Pistil of three fused carpels;
ovary superior
Placentation: parietal or axile
Fruit a capsule
Cyperaceae -> Sedge family
Worldwide: 122 genera, 4500 species.
With approximately 240 species the family is the
third most diverse family in the Louisiana after grasses, Poaceae, (360)
and sunflowers, Asteraceae, (375).
Vegetative features:
Herbs, often rhizomatous
Stems: usually triangular in cross-section, solid,
not jointed.
Leaves: 3-ranked, basal or along lower portion of
the stem
Composed of a blade and a closed sheath, generally
lacking a ligule
Blade linear, grass-like, may be reduced or absent
Reproductive features
Inflorescence a complex arrangement of small spikelets,
usually subtended by bracts.
Flowers bisexual or unisexual, (plants monecious
such as in Carex).
Each flower subtended by a scale-like bract.
Flowers: lacking a perianth (tepals) or composed
of bristles, scales, or hairs
Stamens: 1-3 (-6)
Pistil of 2-3 fused carpels; ovary superior
Placentation: basal
Fruit an achene
Common genera and their recognition features:
Carex: flowers unisexual; monoecious; spikelets
cylindrical and often composed of either pollen bearing
or seed bearing flowers; pistil surrounded by a perigynium;
Scleria: perianth absent, achene white, bony in appearance, without a tubercle.
Fuirena: perianth of 3, stalked, scale-like or paddlelike structures; spikelets with scales spirally attached, usually in terminal umbels.
Elocharis: leaves lacking a blade; aerial stem unbranched, terminated by a single spikelet (spike); achene bearing a persistent tubercle.
Dichromena: Inflorescence subtended by leaflike bracts that are white at the base and green at the tips.
Rhynchospora: spikelets 1-2 flowered; achene bearing a persistent tubercle.
Bulbostylis: typically small plants with leaves capillary or filiform; spikelets many flowered, spiral; achene with a minute, persistent tubercle.
Cyperus: scales of spikelet two-ranked; perianth lacking; spikelets usually in head-like or umbel-like clusters; achenes lacking tubercles.
Cladium: stems 3 meters (ca. 10 ft.) tall;
leaf blades 0.5-1 meter long, margins sharply saw-toothed; spikelets
in clusters on elongate inflorescences; spikelets one-flowered,; flowers
without a perianth.
Fimbristsylis: spikelets several flowered,
scales spirally arranged; spikelets arranged in terminal clusters or simple
to compound umbelliform
cymes; perianth absent; unbranched part of the style is flattened and sometimes
fimbriate; tubercle not persistent.
Scirpus: stems unbranched, triangular to essentially
terete; in some species leaves bladeless; inflorescence variable, without
bracts, with
several leaflike bracts, with a single bract appearing like a continuation
of the stem; spikelets many flowered, scales spirally arranged;
perianth of 1-6 bristles; if style base persistent not tubercle-like.
You may see Scirpus divided into the following genera:
Flora Editorial Committee. 2000. Flora of North America North of Mexico. Vol 22. Magnoliophyta: Alismatidae, Arecidae, Commelinidae (in part), and Zingiberidae. Oxford University Press, New York, etc. [ISBN 0-19-513729-9 (v. 22); LC card number: QK110.F55 2000 581.97 9230459].
Mohlenbrock, R. H. 1999. The Illustrated Flora of Illinois, Sedges: Carex. Southern Illinois University Press, Carbondale and Edwardsville.
Stutzenbaker, C. D. 1999. Aquatic and Wetland Plants
of the Western Gulf Coast. Texas Parks and Wildlife Press. [ISBN 1-885696-31-0].