Lecture three

Grasses and grass-like plants





Grasses and grass-like plants; graminoides.

Poaceae/Gramineae -> grass family

Cyperaceae -> sedge family

Juncaceae -> rush family

Poaceae/Gramineae -> grass family

Contains about 8,500 species in approximatly 650 genera
Ecomomically important
Cereal grains: wheat; corn; rice; barley; rye; etc.
Forage for livestock
Turf for lawns; parks; athletic fields
Source of cover and food for wildlife
Source of construction materials and utensils

Vegetative grass structure
    Roots: adventitious and fibrous
    Stems: specialized types: rhizomes; stolons; runners
    Culm is the upright aerial stem

Recognition Characteristics:
    Jointed with swollen nodes; often hollow at the internodes
    Circular or ellipsoid in cross-section
    Leaves: 2-ranked
    Leaf consists of the blade; open sheath with ligule; auricles may be present
    on the blade; veination is parallel.

Reproductive structure

Inflorescence -> a spikelet which is a two-rankes cluster of florets (small flowers) typically subtended by a pair of glumes (sterile bracts).

Grass flower: adapted for wind or self pollination
    Lemma
    Palea
    Locicules
    Rachilla
    Awns
    Sterile lemma
    Sterile floret
    Stamens: 3 (1-6 or numerous)
    Carpels 3 (often appearing as 2) stigmas 2 (-3) plumose

Grass fruit: a grain or caryopsis

Examples of grasses that typify variation within the family members found in the Gulf Coast Region:

Arundinaria; Hydrochloa; Monanthochloe; Erianthus; Schizachrium; Andropogon; Phragmites; Ctenium; Spartina; Paspalum; Panicum; Zizaniopsis; Setaria; Manisuris; Chinochloa; Eragrostis; Briza; Aristida; Muhlenbergia.
 

Grass identification literature:

Allen, C. 1992. Grasses of Lousiana. Cajun Prairie Habitat Preservation Society. Eunice, LA. [ISBN 0-9633191-0-8; LC card number 92-72282].

Chase, A. 1950. Manual of the Grasses of the United States by A. S. Hetchcock. U.S.D.A. Miscellaneous Publication No. 200. U.S. Government Printing Office, Washington.

Godfrey, R. K. and J. W. Wooten. 1979. Aquatic and Wetland Plants of Southeastern United States; Monocotyledons. University of Georgia Press, Athens. [ISBN: 0-8203-0420-4; LC card number QK125.G6, 584’.0976, 76-28924].

Gould, F. W. 1975. The Grasses of Texas. Texas A & M University Press, College Station. [ISBN 0-89096-005-4, LC card number: QK495.G74G726, 584’.9’097641 75-18688].

Hatch, S. L., J. L. Schuster, and D. L. Drawe. 1999. Grasses of the Texas Gulf Prairies and Marshes. Texas A & M University Press, College Station. [ISBN 0-89096-875-6; LC card number: QK495.G74H34365, 1999, 584’.9’09764-dc21].

Stutzenbaker, C. D. 1999. Aquatic and Wetland Plants of the Western Gulf Coast. Texas Parks and Wildlife Press. [ISBN 1-885696-31-0].
 
 


Juncaceae -> Rush family

Worldwide: 6 genera, 400 species. Louisiana: 2 genera, 30 species.
Juncus and Luzula are two genera found in our region
Juncus: glabrous plants; capsules with many powderlike seeds
Luzula: hairy plants; capsules each with 3 seeds.

Vegetative features:

Herbs
Stems: round and solid, not jointed.
Leaves: 3-ranked, basal or along lower portion of the stem
Composed of a blade and a usually open sheath lacking a ligule
Blade linear, grass-like, may be reduced or absent

Reproductive features

Inflorescence highly branched, but often condensed and head-like
Inflorescence bract may be upright and appear as a continuation of the stem giving the appearance that the flowers are growing from one side of the stem.
i.e.: J. effusus; J. romerianus;

Flowers: 6 distinct perianth (tepals) parts, scale-like.
    Stamens: (3-) 6
    Pistil of three fused carpels; ovary superior

Placentation: parietal or axile
Fruit a capsule

Cyperaceae -> Sedge family

Worldwide: 122 genera, 4500 species.
With approximately 240 species the family is the third most diverse family in the Louisiana after grasses, Poaceae, (360) and sunflowers, Asteraceae, (375).

Vegetative features:

Herbs, often rhizomatous
Stems: usually triangular in cross-section, solid, not jointed.
Leaves: 3-ranked, basal or along lower portion of the stem
Composed of a blade and a closed sheath, generally lacking a ligule
Blade linear, grass-like, may be reduced or absent

Reproductive features

Inflorescence a complex arrangement of small spikelets, usually subtended by bracts.
Flowers bisexual or unisexual, (plants monecious such as in Carex).
Each flower subtended by a scale-like bract.
Flowers: lacking a perianth (tepals) or composed of bristles, scales, or hairs
Stamens: 1-3 (-6)
Pistil of 2-3 fused carpels; ovary superior

Placentation: basal
Fruit an achene

Common genera and their recognition features:

Carex: flowers unisexual; monoecious; spikelets cylindrical and often composed of either pollen bearing
               or seed bearing flowers; pistil surrounded by a perigynium;

Scleria: perianth absent, achene white, bony in appearance, without a tubercle.

Fuirena: perianth of 3, stalked, scale-like or paddlelike structures; spikelets with scales spirally attached, usually in terminal umbels.

Elocharis: leaves lacking a blade; aerial stem unbranched, terminated by a single spikelet (spike); achene bearing a persistent tubercle.

Dichromena: Inflorescence subtended by leaflike bracts that are white at the base and green at the tips.

Rhynchospora: spikelets 1-2 flowered; achene bearing a persistent tubercle.

Bulbostylis: typically small plants with leaves capillary or filiform; spikelets many flowered, spiral; achene with a minute, persistent tubercle.

Cyperus: scales of spikelet two-ranked; perianth lacking; spikelets usually in head-like or umbel-like clusters; achenes lacking tubercles.

Cladium: stems 3 meters (ca. 10 ft.) tall; leaf blades 0.5-1 meter long, margins sharply saw-toothed; spikelets
                  in clusters on elongate inflorescences; spikelets one-flowered,; flowers without a perianth.

Fimbristsylis: spikelets several flowered, scales spirally arranged; spikelets arranged in terminal clusters or simple to compound umbelliform
                         cymes; perianth absent; unbranched part of the style is flattened and sometimes fimbriate; tubercle not persistent.

Scirpus: stems unbranched, triangular to essentially terete; in some species leaves bladeless; inflorescence variable, without bracts, with
                several leaflike bracts, with a single bract appearing like a continuation of the stem; spikelets many flowered, scales spirally arranged;
                perianth of 1-6 bristles; if style base persistent not tubercle-like.

    You may see Scirpus divided into the following genera:

Scirpus americanus Pers. ->Schoenoplectus americanus (Pers.) Volk. ex Schinz & R. Keller; chairmaker's bulrush
Scirpus atrovirens Willd.; green bulrush
Schoenoplectus californicus (C.A. Mey.) Palla, California bulrush, -> Scirpus californicus (C.A. Mey.) Steud.
Oxycaryum cubense (Poepp. & Kunth) Lye, Cuban bulrush, -> Scirpus cubensis Poepp. & Kunth
Scirpus cyperinus (L.) Kunth; woolgrass
Schoenoplectus deltarum (Schuyler) Soják, delta bulrush, ->Scirpus deltarum Schuyler
Scirpus divaricatus Ell.; spreading bulrush
Schoenoplectus etuberculatus (Steud.) Soják, Canby’s bulrush, ->Scirpus etuberculatus (Steud.) Kunze
Isolepis carinata Hook. & Arn. ex Torr., low bulrush,-> Isolepis koilolepis Steud. Scirpus carinatus (Hook. & Arn. ex Torr.)
        Gray, non Sm. Scirpus koilolepis (Steud.) Gleason
Isolepis molesta (M.C. Johnston) S.G. Sm., Gulf Coast bulrush, -> Scirpus molestus M.C. Johnston
Schoenoplectus robustus (Pursh) M.T. Strong; sturdy bulrush, Scirpus robustus Pursh
Schoenoplectus tabernaemontani (K.C. Gmel.) Palla, softstem bulrush; Scirpus validus Vahl
Identification guides: Godfrey, R. K. and J. W. Wooten. 1979. Aquatic and Wetland Plants of Southeastern United States; Monocotyledons. University of Georgia Press, Athens. [ISBN: 0-8203-0420-4; LC card number QK125.G6, 584’.0976, 76-28924].

Flora Editorial Committee. 2000. Flora of North America North of Mexico. Vol 22. Magnoliophyta: Alismatidae, Arecidae, Commelinidae (in part), and Zingiberidae. Oxford University Press, New York, etc. [ISBN 0-19-513729-9 (v. 22); LC card number: QK110.F55 2000 581.97 9230459].

Mohlenbrock, R. H. 1999. The Illustrated Flora of Illinois, Sedges: Carex. Southern Illinois University Press, Carbondale and Edwardsville.

Stutzenbaker, C. D. 1999. Aquatic and Wetland Plants of the Western Gulf Coast. Texas Parks and Wildlife Press. [ISBN 1-885696-31-0].